:root {
  color-scheme: dark;
  --bg: #141c25;
  --panel: #1c2833;
  --panel-alt: #213035;
  --panel-deep: #10171f;
  --text: #f1f1e6;
  --muted: #b8cccb;
  --line: #34464b;
  --line-bright: #486067;
  --accent: #d7e7a8;
  --strong: #67d49b;
  --strong-bg: rgba(103, 212, 155, 0.16);
  --weak: #e8b85c;
  --weak-bg: rgba(232, 184, 92, 0.16);
  --only-a: #68b5e8;
  --danger: #e57373;
  --hatch-pattern: repeating-linear-gradient(
    135deg,
    rgba(20, 28, 37, 0.42) 0px,
    rgba(20, 28, 37, 0.42) 4px,
    transparent 4px,
    transparent 8px
  );
  font-family: Arial, Helvetica, sans-serif;
}

* {
  box-sizing: border-box;
}

body {
  margin: 0;
  background: var(--bg);
  color: var(--text);
  min-height: 100svh;
  font-size: 15px;
  line-height: 1.5;
}

a {
  color: var(--accent);
  text-decoration: none;
  text-underline-offset: 4px;
}

a:hover {
  text-decoration: underline;
}

button,
input,
select {
  font: inherit;
  color: inherit;
}

button:focus-visible,
a:focus-visible,
input:focus-visible,
select:focus-visible,
[tabindex]:focus-visible {
  outline: 3px solid var(--accent);
  outline-offset: 3px;
}

[hidden] {
  display: none !important;
}

.site-header,
main,
footer {
  width: min(1540px, calc(100% - 48px));
  margin-inline: auto;
}

.site-header {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 20px;
  min-height: 86px;
  border-bottom: 1px solid var(--line);
}

.brand {
  font: 700 33px Georgia, 'Times New Roman', serif;
  letter-spacing: -1px;
  color: var(--text);
  text-decoration: none;
}

.brand:hover {
  text-decoration: none;
}

.brand span {
  font: 400 13px Arial, Helvetica, sans-serif;
  letter-spacing: 1.5px;
  margin-left: 10px;
  color: var(--muted);
}

.site-header nav {
  display: flex;
  align-items: center;
  gap: 24px;
  color: var(--muted);
  font-size: 15px;
}

.site-header nav a {
  color: var(--muted);
}

.site-header nav a:hover {
  color: var(--text);
}

footer {
  border-top: 1px solid var(--line);
  display: flex;
  justify-content: space-between;
  align-items: center;
  gap: 20px;
  padding-block: 24px;
  margin-top: 44px;
  font-size: 14px;
  color: var(--muted);
}

/* Cross-navigation bar for all 7 genomics tools */
.gene-tools-nav {
  display: flex;
  flex-wrap: wrap;
  gap: 8px;
  padding-block: 14px;
  border-bottom: 1px solid var(--line);
}

.gene-tool-link {
  display: inline-flex;
  align-items: center;
  padding: 6px 12px;
  border-radius: 6px;
  background: var(--panel);
  border: 1px solid var(--line);
  color: var(--muted);
  font-size: 13px;
  font-weight: 600;
  text-decoration: none;
  transition: background 0.12s, border-color 0.12s, color 0.12s;
}

.gene-tool-link:hover {
  background: var(--panel-alt);
  border-color: var(--line-bright);
  color: var(--text);
  text-decoration: none;
}

.gene-tool-link.active {
  background: rgba(215, 231, 168, 0.14);
  border-color: var(--accent);
  color: var(--accent);
}

/* Page Top Heading */
.page-top {
  display: flex;
  justify-content: space-between;
  align-items: flex-start;
  gap: 28px;
  padding-block: 24px 20px;
  border-bottom: 1px solid var(--line);
  margin-bottom: 20px;
}

.eyebrow-row {
  display: flex;
  align-items: center;
  flex-wrap: wrap;
  gap: 10px;
  margin-bottom: 8px;
}

.meta-pill {
  font-size: 12.5px;
  color: var(--muted);
  font-variant-numeric: tabular-nums;
}

.page-top h1 {
  font: 600 clamp(26px, 2.5vw, 35px)/1.2 Georgia, serif;
  margin: 0 0 8px;
  color: var(--text);
}

.page-top .lead {
  color: var(--muted);
  font-size: 15px;
  max-width: 860px;
  margin: 0;
}

.lead-compact {
  color: var(--muted);
  font-size: 14px;
  margin: 6px 0 0;
}

.tier-legend-box {
  display: grid;
  gap: 8px;
  background: var(--panel);
  border: 1px solid var(--line);
  border-radius: 7px;
  padding: 12px 15px;
  min-width: 310px;
  max-width: 410px;
  flex-shrink: 0;
}

.legend-item {
  display: flex;
  align-items: center;
  gap: 10px;
  font-size: 12.5px;
}

.legend-item div {
  display: flex;
  flex-direction: column;
  line-height: 1.3;
}

.legend-item strong {
  color: var(--text);
}

.legend-item span {
  color: var(--muted);
  font-size: 11.5px;
}

.swatch {
  width: 22px;
  height: 14px;
  border-radius: 3px;
  flex-shrink: 0;
  border: 1px solid rgba(255, 255, 255, 0.2);
}

.swatch-strong {
  background: var(--strong);
}

.swatch-weak {
  background: var(--weak);
}

.swatch-absent {
  background: #23313a;
}

/* Loading Banner */
.loading-panel {
  display: flex;
  align-items: center;
  gap: 18px;
  background: var(--panel);
  border: 1px solid var(--line);
  border-radius: 8px;
  padding: 24px 28px;
  margin-block: 20px;
}

.loading-panel p {
  margin: 4px 0 0;
  color: var(--muted);
  font-size: 14px;
}

.spinner {
  width: 28px;
  height: 28px;
  border-radius: 50%;
  border: 3px solid var(--line);
  border-top-color: var(--accent);
  animation: spin 0.85s linear infinite;
  flex-shrink: 0;
}

@keyframes spin {
  to {
    transform: rotate(360deg);
  }
}

/* Panels */
.panel {
  background: var(--panel);
  border: 1px solid var(--line);
  border-radius: 8px;
  padding: 20px 22px;
  margin-bottom: 20px;
}

.section-head {
  display: flex;
  justify-content: space-between;
  align-items: flex-start;
  gap: 16px;
  margin-bottom: 16px;
}

.section-head h2 {
  font: 600 21px/1.25 Georgia, serif;
  margin: 2px 0 0;
  color: var(--text);
}

.section-head p {
  margin: 4px 0 0;
  color: var(--muted);
  font-size: 13.5px;
}

.step-tag {
  display: inline-block;
  font-size: 11px;
  font-weight: 700;
  text-transform: uppercase;
  letter-spacing: 0.7px;
  color: var(--accent);
}

.count-pill,
.compute-badge {
  display: inline-flex;
  align-items: center;
  padding: 4px 10px;
  border-radius: 999px;
  background: var(--panel-alt);
  border: 1px solid var(--line);
  color: var(--accent);
  font-size: 12px;
  font-weight: 600;
  font-variant-numeric: tabular-nums;
  white-space: nowrap;
}

.control-field {
  display: grid;
  gap: 5px;
  font-size: 12.5px;
  color: var(--muted);
  min-width: 0;
}

.control-field.inline-field {
  min-width: 300px;
}

.control-field > span {
  font-weight: 600;
  color: #d2e0df;
}

input[type="search"],
input[type="text"],
select {
  width: 100%;
  min-height: 38px;
  padding: 7px 11px;
  border: 1px solid var(--line-bright);
  border-radius: 5px;
  background: var(--panel-deep);
  color: var(--text);
  font-size: 14px;
}

.creature-select {
  font-weight: 600;
  min-height: 38px;
  background: #131d26;
  border-color: #5b7880;
}

/* 10 Trait Modules Grid */
.trait-modules-grid {
  display: grid;
  grid-template-columns: repeat(5, minmax(0, 1fr));
  gap: 12px;
}

.trait-card {
  cursor: pointer;
  text-align: left;
  background: var(--panel-deep);
  border: 1px solid var(--line);
  border-radius: 7px;
  padding: 13px 14px;
  display: flex;
  flex-direction: column;
  justify-content: space-between;
  gap: 10px;
  transition: border-color 0.12s, background 0.12s, transform 0.1s;
}

.trait-card:hover {
  border-color: var(--line-bright);
  background: var(--panel-alt);
  transform: translateY(-1px);
}

.trait-card.active {
  background: rgba(215, 231, 168, 0.1);
  border-color: var(--accent);
  box-shadow: inset 0 0 0 1px var(--accent);
}

.trait-card-title {
  font: 600 14.5px/1.25 Georgia, serif;
  color: var(--text);
  margin: 4px 0 0;
}

.trait-card.active .trait-card-title {
  color: var(--accent);
}

.trait-card-sub {
  font-size: 11.5px;
  color: var(--muted);
  line-height: 1.35;
}

.trait-card-foot {
  display: flex;
  justify-content: space-between;
  align-items: center;
  padding-top: 7px;
  border-top: 1px solid #23333b;
  font-size: 11.5px;
  color: var(--muted);
}

.trait-card-foot strong {
  color: var(--text);
}

/* Trait Highlight Box */
.trait-highlight-top {
  display: grid;
  grid-template-columns: 1.4fr 1fr;
  gap: 20px;
  align-items: start;
  padding-bottom: 16px;
  border-bottom: 1px solid var(--line);
}

.trait-highlight-top h2 {
  font: 600 24px/1.25 Georgia, serif;
  margin: 4px 0 0;
}

.trait-kpi-strip {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 10px;
}

.mini-kpi-box {
  background: var(--panel-deep);
  border: 1px solid var(--line);
  border-radius: 6px;
  padding: 10px 12px;
}

.mini-kpi-box span {
  display: block;
  font-size: 11px;
  font-weight: 700;
  text-transform: uppercase;
  color: var(--muted);
}

.mini-kpi-box strong {
  display: block;
  font-size: 20px;
  margin-top: 3px;
  color: var(--accent);
  font-variant-numeric: tabular-nums;
}

.mini-kpi-box small {
  font-size: 11px;
  color: var(--muted);
}

.core-genes-roles-section {
  margin-top: 14px;
}

.core-genes-roles-grid {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 9px;
  margin-top: 8px;
}

.core-gene-chip {
  cursor: pointer;
  text-align: left;
  background: var(--panel-deep);
  border: 1px solid var(--line);
  border-radius: 6px;
  padding: 8px 10px;
  display: flex;
  flex-direction: column;
  gap: 3px;
  transition: border-color 0.12s, background 0.12s;
}

.core-gene-chip:hover {
  border-color: var(--accent);
  background: var(--panel-alt);
}

.core-gene-chip-top {
  display: flex;
  justify-content: space-between;
  align-items: baseline;
  gap: 6px;
}

.core-gene-chip-top strong {
  color: var(--accent);
  font-size: 13px;
}

.core-gene-chip-top small {
  color: var(--muted);
  font-size: 11px;
  font-variant-numeric: tabular-nums;
}

.core-gene-chip-role {
  font-size: 11.5px;
  color: var(--text);
  line-height: 1.3;
}

/* Species x Trait-Gene Heatmap Matrix */
.matrix-add-species-bar {
  display: grid;
  grid-template-columns: 1.3fr 1.4fr auto;
  gap: 12px;
  align-items: end;
  padding: 13px 15px;
  background: var(--panel-deep);
  border: 1px solid var(--line);
  border-radius: 7px;
  margin-bottom: 14px;
}

.matrix-scroll {
  width: 100%;
  overflow: auto;
  max-height: 680px;
  border: 1px solid var(--line);
  border-radius: 7px;
  background: var(--panel-deep);
}

.trait-heatmap-table {
  border-collapse: separate;
  border-spacing: 2px;
  font-size: 12px;
  width: max-content;
  min-width: 100%;
}

.trait-heatmap-table th,
.trait-heatmap-table td {
  padding: 6px 8px;
  font-variant-numeric: tabular-nums;
}

.trait-heatmap-table thead th {
  position: sticky;
  top: 0;
  z-index: 3;
  background: #141e27;
  color: var(--accent);
  font-weight: 700;
  border-bottom: 1px solid var(--line-bright);
  text-align: center;
  white-space: nowrap;
}

.trait-heatmap-table thead th.corner-th {
  position: sticky;
  left: 0;
  z-index: 4;
  text-align: left;
  min-width: 240px;
  color: var(--text);
}

.trait-heatmap-table tbody th {
  position: sticky;
  left: 0;
  z-index: 2;
  background: #141e27;
  color: var(--text);
  text-align: left;
  white-space: nowrap;
  border-right: 1px solid var(--line);
}

.hm-row-head {
  display: flex;
  justify-content: space-between;
  align-items: center;
  gap: 8px;
}

.hm-remove-row-btn {
  cursor: pointer;
  background: transparent;
  border: 0;
  color: var(--muted);
  font-size: 11px;
  padding: 2px 5px;
}

.hm-remove-row-btn:hover {
  color: var(--danger);
}

.hm-cell {
  cursor: pointer;
  text-align: center;
  font-size: 10.5px;
  font-weight: 700;
  border-radius: 3px;
  min-width: 58px;
}

.hm-cell.strong {
  background: rgba(103, 212, 155, 0.25);
  color: var(--strong);
  border: 1px solid rgba(103, 212, 155, 0.45);
}

.hm-cell.weak {
  background: rgba(232, 184, 92, 0.24);
  color: var(--weak);
  border: 1px solid rgba(232, 184, 92, 0.45);
}

.hm-cell.absent {
  background: rgba(255, 255, 255, 0.03);
  color: #5d747c;
  border: 1px solid #223038;
}

/* Data Tables & Filters */
.table-filters-bar {
  display: grid;
  grid-template-columns: 1.5fr 1fr 1.15fr 1.15fr 1.15fr;
  gap: 12px;
  padding: 14px 16px;
  background: var(--panel-deep);
  border: 1px solid var(--line);
  border-radius: 7px;
  margin-bottom: 14px;
}

.table-filters-bar.filters-bar-2 {
  grid-template-columns: 1.6fr 1fr;
}

.table-scroll {
  width: 100%;
  overflow-x: auto;
  border: 1px solid var(--line);
  border-radius: 7px;
  background: var(--panel-deep);
}

.data-table {
  width: 100%;
  border-collapse: collapse;
  font-size: 13.5px;
  text-align: left;
}

.data-table th,
.data-table td {
  padding: 10px 12px;
  border-bottom: 1px solid #25343c;
  vertical-align: middle;
}

.data-table thead th {
  position: sticky;
  top: 0;
  z-index: 2;
  background: #17222b;
  color: #cfe0df;
  font-size: 12px;
  font-weight: 700;
  text-transform: uppercase;
  letter-spacing: 0.4px;
  border-bottom: 1px solid var(--line-bright);
  white-space: nowrap;
}

.data-table tbody tr:hover {
  background: rgba(215, 231, 168, 0.05);
}

.clickable-row {
  cursor: pointer;
}

.col-rank {
  width: 54px;
  color: var(--muted);
  font-variant-numeric: tabular-nums;
}

.col-num {
  text-align: right;
  font-variant-numeric: tabular-nums;
  white-space: nowrap;
}

.col-bar {
  min-width: 185px;
  width: 215px;
}

.col-action {
  text-align: right;
  white-space: nowrap;
}

.num-strong {
  color: var(--strong);
  font-weight: 700;
}

.num-weak {
  color: var(--weak);
  font-weight: 700;
}

.stacked-bar-track {
  width: 100%;
  height: 10px;
  background: #0d1319;
  border: 1px solid var(--line);
  border-radius: 999px;
  overflow: hidden;
  display: flex;
}

.bar-seg {
  height: 100%;
}

.bar-strong {
  background-color: var(--strong);
}

.bar-weak {
  background-color: var(--weak);
}

.badge-row {
  display: flex;
  flex-wrap: wrap;
  gap: 7px;
  margin-bottom: 6px;
}

.badge {
  display: inline-flex;
  align-items: center;
  padding: 3px 9px;
  border-radius: 4px;
  font-size: 11.5px;
  font-weight: 700;
  border: 1px solid var(--line);
  background: var(--panel-alt);
  color: var(--text);
}

.badge-group {
  background: rgba(215, 231, 168, 0.12);
  border-color: rgba(215, 231, 168, 0.35);
  color: var(--accent);
}

.badge-kingdom {
  background: rgba(104, 181, 232, 0.14);
  border-color: rgba(104, 181, 232, 0.4);
  color: #9cd2f5;
}

.badge-consensus {
  background: rgba(103, 212, 155, 0.15);
  border-color: rgba(103, 212, 155, 0.45);
  color: var(--strong);
}

.badge-provisional {
  background-color: rgba(232, 184, 92, 0.16);
  background-image: var(--hatch-pattern);
  border: 1px dashed rgba(232, 184, 92, 0.65);
  color: var(--weak);
}

.btn-accent,
.btn-secondary,
.btn-inline {
  cursor: pointer;
  border-radius: 5px;
  font-weight: 700;
  font-size: 13px;
  display: inline-flex;
  align-items: center;
  justify-content: center;
  gap: 6px;
  white-space: nowrap;
  transition: background 0.12s, border-color 0.12s;
}

.btn-accent {
  background: var(--accent);
  color: #162225;
  border: 1px solid var(--accent);
  padding: 8px 14px;
  min-height: 38px;
}

.btn-secondary {
  background: var(--panel-alt);
  color: var(--text);
  border: 1px solid var(--line-bright);
  padding: 8px 13px;
  min-height: 38px;
}

.btn-secondary:hover:not(:disabled) {
  background: #2c4047;
  border-color: var(--accent);
}

.btn-secondary:disabled {
  opacity: 0.45;
  cursor: not-allowed;
}

.btn-inline {
  background: rgba(215, 231, 168, 0.14);
  color: var(--accent);
  border: 1px solid rgba(215, 231, 168, 0.4);
  padding: 4px 9px;
  font-size: 11.5px;
}

.btn-inline:hover {
  background: var(--accent);
  color: #162225;
}

.pagination-bar {
  display: flex;
  justify-content: space-between;
  align-items: center;
  flex-wrap: wrap;
  gap: 12px;
  margin-top: 14px;
  font-size: 13px;
  color: var(--muted);
}

.page-buttons {
  display: flex;
  gap: 8px;
}

/* Modal */
.modal-backdrop {
  position: fixed;
  inset: 0;
  z-index: 100;
  background: rgba(9, 14, 19, 0.82);
  display: flex;
  align-items: center;
  justify-content: center;
  padding: 20px;
}

.modal-dialog {
  width: min(980px, 100%);
  max-height: calc(100vh - 40px);
  background: var(--panel);
  border: 1px solid var(--line-bright);
  border-radius: 9px;
  display: flex;
  flex-direction: column;
  overflow: hidden;
  box-shadow: 0 20px 60px rgba(0, 0, 0, 0.65);
}

.modal-header {
  display: flex;
  justify-content: space-between;
  align-items: flex-start;
  gap: 16px;
  padding: 18px 22px;
  background: var(--panel-deep);
  border-bottom: 1px solid var(--line);
}

.modal-header h2 {
  font: 600 22px/1.25 Georgia, serif;
  margin: 0;
}

.modal-sub {
  margin: 4px 0 0;
  color: var(--muted);
  font-size: 13.5px;
}

.modal-close-btn {
  cursor: pointer;
  padding: 8px 14px;
  border-radius: 5px;
  border: 1px solid var(--line-bright);
  background: var(--panel-alt);
  color: var(--text);
  font-weight: 700;
}

.modal-body {
  padding: 18px 22px;
  overflow-y: auto;
  display: grid;
  gap: 16px;
}

.modal-card {
  background: var(--panel-deep);
  border: 1px solid var(--line);
  border-radius: 7px;
  padding: 14px 16px;
}

.modal-card h3 {
  font: 600 16px/1.25 Georgia, serif;
  margin: 0 0 10px;
}

.heatmap-modal-strip {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 8px;
}

@media (max-width: 1240px) {
  .trait-modules-grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }
  .trait-highlight-top,
  .matrix-add-species-bar {
    grid-template-columns: 1fr;
  }
  .core-genes-roles-grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }
}

@media (max-width: 780px) {
  .site-header,
  main,
  footer {
    width: calc(100% - 28px);
  }
  .page-top {
    flex-direction: column;
  }
  .tier-legend-box {
    width: 100%;
    max-width: none;
  }
  .trait-modules-grid,
  .core-genes-roles-grid,
  .trait-kpi-strip,
  .table-filters-bar,
  .table-filters-bar.filters-bar-2,
  .heatmap-modal-strip {
    grid-template-columns: 1fr;
  }
}