:root {
  color-scheme: dark;
  --bg: #141c25;
  --panel: #1c2833;
  --panel-alt: #213035;
  --panel-deep: #0e151d;
  --text: #f1f1e6;
  --muted: #b8cccb;
  --line: #34464b;
  --line-bright: #4b656d;
  --accent: #d7e7a8;
  --strong: #67d49b;
  --weak: #e8b85c;
  --focal: #5ad2ff;
  --danger: #ef767a;
  font-family: Arial, Helvetica, sans-serif;
}

* {
  box-sizing: border-box;
}

body {
  margin: 0;
  background: var(--bg);
  color: var(--text);
  min-height: 100svh;
  font-size: 14px;
  line-height: 1.45;
}

a {
  color: var(--accent);
  text-decoration: none;
}

a:hover {
  text-decoration: underline;
}

button,
input,
select {
  font: inherit;
  color: inherit;
}

[hidden] {
  display: none !important;
}

.site-header,
main,
footer {
  width: min(1680px, calc(100% - 36px));
  margin-inline: auto;
}

.site-header {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 20px;
  min-height: 78px;
  border-bottom: 1px solid var(--line);
}

.brand {
  font: 700 31px Georgia, 'Times New Roman', serif;
  letter-spacing: -1px;
  color: var(--text);
  text-decoration: none;
}

.brand span {
  font: 400 13px Arial, Helvetica, sans-serif;
  letter-spacing: 1.5px;
  margin-left: 10px;
  color: var(--muted);
}

.site-header nav {
  display: flex;
  align-items: center;
  gap: 22px;
  color: var(--muted);
  font-size: 15px;
}

.site-header nav a {
  color: var(--muted);
}

.site-header nav a:hover {
  color: var(--text);
}

footer {
  border-top: 1px solid var(--line);
  display: flex;
  justify-content: space-between;
  align-items: center;
  padding-block: 20px;
  margin-top: 32px;
  color: var(--muted);
  font-size: 13.5px;
}

/* Cross-navigation bar */
.gene-tools-nav {
  display: flex;
  flex-wrap: wrap;
  gap: 7px;
  padding-block: 12px;
  border-bottom: 1px solid var(--line);
}

.gene-tool-link {
  display: inline-flex;
  align-items: center;
  padding: 5px 11px;
  border-radius: 6px;
  background: var(--panel);
  border: 1px solid var(--line);
  color: var(--muted);
  font-size: 12.5px;
  font-weight: 600;
  text-decoration: none;
  transition: background 0.12s, border-color 0.12s, color 0.12s;
}

.gene-tool-link:hover {
  background: var(--panel-alt);
  border-color: var(--line-bright);
  color: var(--text);
  text-decoration: none;
}

.gene-tool-link.active {
  background: rgba(215, 231, 168, 0.15);
  border-color: var(--accent);
  color: var(--accent);
}

/* Page Top */
.page-top {
  display: flex;
  justify-content: space-between;
  align-items: flex-start;
  gap: 20px;
  padding-block: 18px 14px;
}

.meta-pill {
  font-size: 12px;
  color: var(--accent);
  font-weight: 600;
}

.page-top h1 {
  font: 600 clamp(24px, 2.2vw, 32px)/1.2 Georgia, serif;
  margin: 4px 0 6px;
}

.page-top .lead {
  margin: 0;
  color: var(--muted);
  max-width: 920px;
  font-size: 14px;
}

.toolbar-actions {
  display: flex;
  gap: 10px;
  flex-shrink: 0;
}

.btn-accent,
.btn-secondary,
.btn-inline,
.btn-tiny {
  cursor: pointer;
  border-radius: 6px;
  font-weight: 700;
  transition: background 0.12s, border-color 0.12s, transform 0.08s;
}

.btn-accent {
  background: var(--accent);
  color: #142024;
  border: 1px solid var(--accent);
  padding: 8px 14px;
  font-size: 13px;
}

.btn-accent:hover {
  background: #e4f2be;
}

.btn-secondary {
  background: var(--panel);
  color: var(--text);
  border: 1px solid var(--line-bright);
  padding: 8px 13px;
  font-size: 13px;
}

.btn-secondary:hover {
  border-color: var(--accent);
}

.btn-inline {
  background: rgba(215, 231, 168, 0.14);
  color: var(--accent);
  border: 1px solid rgba(215, 231, 168, 0.4);
  padding: 4px 10px;
  font-size: 12px;
}

.btn-inline:hover {
  background: var(--accent);
  color: #142024;
}

.btn-tiny {
  background: var(--panel-deep);
  color: var(--muted);
  border: 1px solid var(--line);
  padding: 2px 8px;
  font-size: 11px;
}

.btn-tiny:hover {
  color: var(--accent);
  border-color: var(--accent);
}

/* Loading Panel */
.loading-panel {
  display: flex;
  align-items: center;
  gap: 16px;
  background: var(--panel);
  border: 1px solid var(--line);
  border-radius: 8px;
  padding: 22px 26px;
  margin-block: 16px;
}

.spinner {
  width: 26px;
  height: 26px;
  border-radius: 50%;
  border: 3px solid var(--line);
  border-top-color: var(--accent);
  animation: spin 0.85s linear infinite;
}

@keyframes spin {
  to {
    transform: rotate(360deg);
  }
}

/* Mode Bar & Evolutionary Slider */
.panel {
  background: var(--panel);
  border: 1px solid var(--line);
  border-radius: 8px;
  padding: 14px 16px;
  margin-bottom: 14px;
}

.scene-modes-row {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 10px;
  margin-bottom: 12px;
}

.scene-mode-btn {
  cursor: pointer;
  text-align: left;
  background: var(--panel-deep);
  border: 1px solid var(--line);
  border-radius: 7px;
  padding: 10px 14px;
  display: flex;
  flex-direction: column;
  gap: 3px;
  transition: border-color 0.12s, background 0.12s;
}

.scene-mode-btn:hover {
  border-color: var(--line-bright);
  background: var(--panel-alt);
}

.scene-mode-btn.active {
  background: rgba(215, 231, 168, 0.12);
  border-color: var(--accent);
  box-shadow: inset 0 0 0 1px var(--accent);
}

.scene-mode-btn strong {
  font-size: 14px;
  color: var(--text);
}

.scene-mode-btn.active strong {
  color: var(--accent);
}

.scene-mode-btn span {
  font-size: 11.5px;
  color: var(--muted);
}

.evo-slider-bar {
  background: var(--panel-deep);
  border: 1px solid var(--line);
  border-radius: 7px;
  padding: 10px 14px;
}

.evo-slider-head {
  display: flex;
  justify-content: space-between;
  align-items: center;
  gap: 12px;
  margin-bottom: 6px;
  font-size: 12.5px;
}

.evo-slider-controls {
  display: flex;
  gap: 8px;
}

#evoTimelineSlider {
  width: 100%;
  accent-color: var(--accent);
  cursor: pointer;
  margin: 4px 0 6px;
}

.evo-stops-row {
  display: grid;
  grid-template-columns: repeat(16, minmax(0, 1fr));
  gap: 4px;
}

.evo-stop-btn {
  cursor: pointer;
  background: #141e27;
  border: 1px solid #263740;
  border-radius: 4px;
  padding: 4px 3px;
  font-size: 10.5px;
  color: var(--muted);
  text-align: center;
  white-space: nowrap;
  overflow: hidden;
  text-overflow: ellipsis;
}

.evo-stop-btn:hover,
.evo-stop-btn.active {
  border-color: var(--accent);
  color: var(--accent);
  background: rgba(215, 231, 168, 0.14);
  font-weight: 700;
}

/* 3-Column Studio Grid */
.studio-grid {
  display: grid;
  grid-template-columns: 330px minmax(0, 1fr) 360px;
  gap: 14px;
  align-items: start;
}

.studio-sidebar {
  display: flex;
  flex-direction: column;
  gap: 12px;
}

.sidebar-block {
  background: var(--panel);
  border: 1px solid var(--line);
  border-radius: 8px;
  padding: 12px 13px;
}

.block-head {
  display: flex;
  justify-content: space-between;
  align-items: baseline;
  gap: 8px;
  margin-bottom: 8px;
}

.block-head h3 {
  font: 600 14.5px/1.2 Georgia, serif;
  margin: 0;
  color: var(--text);
}

.block-head small {
  font-size: 11px;
  color: var(--muted);
}

.search-label {
  display: block;
  font-size: 11.5px;
  font-weight: 700;
  color: var(--accent);
  margin-bottom: 6px;
}

input[type="search"] {
  width: 100%;
  padding: 7px 10px;
  border-radius: 5px;
  border: 1px solid var(--line-bright);
  background: var(--panel-deep);
  color: var(--text);
  font-size: 13px;
}

/* Hover-Responsive Pill Lists & Grids */
.hover-pill-list {
  display: flex;
  flex-direction: column;
  gap: 4px;
  margin-top: 7px;
  max-height: 175px;
  overflow-y: auto;
}

.hover-pill-item {
  cursor: pointer;
  display: flex;
  justify-content: space-between;
  align-items: center;
  padding: 5px 8px;
  border-radius: 5px;
  background: var(--panel-deep);
  border: 1px solid #24343d;
  font-size: 12px;
  text-align: left;
}

.hover-pill-item:hover,
.hover-pill-item.active {
  border-color: var(--accent);
  background: rgba(215, 231, 168, 0.12);
}

.splits-pill-grid {
  display: grid;
  grid-template-columns: 1fr;
  gap: 5px;
  max-height: 235px;
  overflow-y: auto;
  padding-right: 2px;
}

.split-btn {
  cursor: pointer;
  text-align: left;
  background: var(--panel-deep);
  border: 1px solid #263740;
  border-radius: 6px;
  padding: 6px 9px;
  display: flex;
  flex-direction: column;
  gap: 2px;
  transition: border-color 0.1s, background 0.1s;
}

.split-btn:hover,
.split-btn.active {
  border-color: var(--accent);
  background: rgba(215, 231, 168, 0.13);
}

.split-btn strong {
  font-size: 12px;
  color: var(--accent);
}

.split-btn span {
  font-size: 11px;
  color: var(--muted);
}

.hover-pill-grid {
  display: flex;
  flex-wrap: wrap;
  gap: 5px;
}

.groups-scroll {
  max-height: 220px;
  overflow-y: auto;
  padding-right: 2px;
}

.clade-pill {
  cursor: pointer;
  background: var(--panel-deep);
  border: 1px solid #283a43;
  border-radius: 5px;
  padding: 4px 8px;
  font-size: 11.5px;
  color: var(--text);
  display: inline-flex;
  align-items: center;
  gap: 5px;
}

.clade-pill small {
  color: var(--muted);
  font-size: 10.5px;
}

.clade-pill:hover,
.clade-pill.active {
  border-color: var(--accent);
  background: rgba(215, 231, 168, 0.16);
  color: var(--accent);
}

/* Center 3D Viewport */
.viewport-wrapper {
  background: var(--panel);
  border: 1px solid var(--line);
  border-radius: 8px;
  padding: 10px 12px;
  display: flex;
  flex-direction: column;
  gap: 10px;
}

.viewport-top-bar {
  display: flex;
  justify-content: space-between;
  align-items: center;
  flex-wrap: wrap;
  gap: 10px;
  padding: 4px 6px;
}

.viewport-status-left {
  display: flex;
  align-items: center;
  flex-wrap: wrap;
  gap: 8px;
}

.active-focal-tag {
  font-size: 11.5px;
  text-transform: uppercase;
  letter-spacing: 0.6px;
  color: var(--muted);
  font-weight: 700;
}

.badge-row-inline {
  display: inline-flex;
  gap: 6px;
}

.badge {
  display: inline-flex;
  align-items: center;
  padding: 2px 8px;
  border-radius: 4px;
  font-size: 11px;
  font-weight: 700;
  background: var(--panel-deep);
  border: 1px solid var(--line);
  color: var(--accent);
}

.viewport-toggles {
  display: flex;
  gap: 12px;
  font-size: 12px;
  color: var(--muted);
}

.toggle-chip {
  display: inline-flex;
  align-items: center;
  gap: 5px;
  cursor: pointer;
}

.canvas-container {
  position: relative;
  width: 100%;
  height: 620px;
  border-radius: 7px;
  overflow: hidden;
  border: 1px solid var(--line-bright);
  background: radial-gradient(circle at 50% 42%, #182734 0%, #0b1219 75%, #070c11 100%);
}

#threeCanvas {
  width: 100%;
  height: 100%;
  display: block;
  cursor: grab;
}

#threeCanvas:active {
  cursor: grabbing;
}

/* Floating 3D Hover Tooltip */
.hover-tooltip {
  position: absolute;
  pointer-events: none;
  z-index: 20;
  min-width: 230px;
  max-width: 310px;
  background: rgba(12, 19, 26, 0.95);
  border: 1px solid var(--accent);
  border-radius: 7px;
  padding: 9px 11px;
  box-shadow: 0 10px 28px rgba(0, 0, 0, 0.65);
  font-size: 12px;
  line-height: 1.35;
}

.hover-tooltip strong {
  color: var(--accent);
  font-size: 13.5px;
  display: block;
}

.hover-tooltip .tt-sub {
  color: var(--muted);
  font-size: 11px;
  margin-bottom: 5px;
}

.hover-tooltip .tt-metrics {
  display: flex;
  justify-content: space-between;
  gap: 8px;
  padding-top: 5px;
  border-top: 1px solid #263842;
  font-variant-numeric: tabular-nums;
}

.viewport-hud-bottom {
  position: absolute;
  left: 10px;
  right: 10px;
  bottom: 8px;
  display: flex;
  justify-content: space-between;
  align-items: center;
  flex-wrap: wrap;
  gap: 8px;
  pointer-events: none;
  font-size: 11px;
  color: var(--muted);
  background: rgba(11, 18, 25, 0.78);
  border: 1px solid rgba(75, 101, 109, 0.45);
  border-radius: 5px;
  padding: 5px 10px;
}

.hud-legend {
  display: flex;
  flex-wrap: wrap;
  gap: 12px;
}

.dot {
  display: inline-block;
  width: 9px;
  height: 9px;
  border-radius: 50%;
  margin-right: 4px;
  vertical-align: middle;
}

.dot-focal {
  background: var(--focal);
  box-shadow: 0 0 6px var(--focal);
}

.dot-strong {
  background: var(--strong);
}

.dot-mid {
  background: var(--weak);
}

.dot-low {
  background: #6c8796;
}

/* 16 Organ Hover Strip */
.organ-hover-strip-box {
  background: var(--panel-deep);
  border: 1px solid var(--line);
  border-radius: 6px;
  padding: 8px 10px;
}

.block-head-inline {
  display: flex;
  justify-content: space-between;
  align-items: center;
  margin-bottom: 6px;
  font-size: 12px;
}

.organ-hover-strip {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 5px;
}

.organ-pill {
  cursor: pointer;
  background: #141f28;
  border: 1px solid #263842;
  border-radius: 5px;
  padding: 5px 7px;
  text-align: left;
  display: flex;
  justify-content: space-between;
  align-items: center;
  gap: 4px;
  font-size: 11px;
}

.organ-pill:hover,
.organ-pill.active {
  border-color: var(--accent);
  background: rgba(215, 231, 168, 0.15);
}

.organ-pill span {
  overflow: hidden;
  text-overflow: ellipsis;
  white-space: nowrap;
}

.organ-pill strong {
  color: var(--strong);
  font-variant-numeric: tabular-nums;
}

/* Right Inspector */
.eyebrow-small {
  font-size: 10.5px;
  font-weight: 700;
  letter-spacing: 0.7px;
  color: var(--accent);
}

.focal-summary-card h2 {
  font: 600 21px/1.2 Georgia, serif;
  margin: 2px 0;
}

.sci-sub {
  margin: 0 0 10px;
  color: var(--muted);
  font-size: 12.5px;
}

.inspector-kpi-grid {
  display: grid;
  grid-template-columns: repeat(2, 1fr);
  gap: 7px;
}

.ikpi {
  background: var(--panel-deep);
  border: 1px solid var(--line);
  border-radius: 5px;
  padding: 7px 9px;
}

.ikpi span {
  display: block;
  font-size: 10.5px;
  color: var(--muted);
  text-transform: uppercase;
}

.ikpi strong {
  font-size: 16px;
  color: var(--text);
  font-variant-numeric: tabular-nums;
}

.neighbors-list {
  display: flex;
  flex-direction: column;
  gap: 4px;
  max-height: 225px;
  overflow-y: auto;
}

.neighbor-row {
  cursor: pointer;
  display: flex;
  justify-content: space-between;
  align-items: center;
  padding: 5px 8px;
  border-radius: 5px;
  background: var(--panel-deep);
  border: 1px solid #23333c;
  font-size: 12px;
}

.neighbor-row:hover {
  border-color: var(--focal);
  background: rgba(90, 210, 255, 0.12);
}

.mini-tabs {
  display: flex;
  gap: 4px;
}

.mini-tab {
  cursor: pointer;
  padding: 3px 7px;
  border-radius: 4px;
  border: 1px solid var(--line);
  background: var(--panel-deep);
  font-size: 11px;
  color: var(--muted);
}

.mini-tab.active {
  border-color: var(--accent);
  color: var(--accent);
  background: rgba(215, 231, 168, 0.14);
}

.context-note {
  margin: 0 0 7px;
  font-size: 11.5px;
  color: var(--muted);
}

.inspector-genes-list {
  margin-top: 8px;
  display: flex;
  flex-direction: column;
  gap: 5px;
  max-height: 330px;
  overflow-y: auto;
}

.igene-card {
  display: flex;
  justify-content: space-between;
  align-items: center;
  gap: 8px;
  padding: 6px 9px;
  border-radius: 5px;
  background: var(--panel-deep);
  border: 1px solid #23333c;
  font-size: 12px;
}

.igene-card:hover {
  border-color: var(--accent);
}

.igene-card strong {
  color: var(--accent);
}

@media (max-width: 1320px) {
  .studio-grid {
    grid-template-columns: 1fr;
  }
  .evo-stops-row {
    grid-template-columns: repeat(8, minmax(0, 1fr));
  }
}

@media (max-width: 780px) {
  .scene-modes-row,
  .organ-hover-strip {
    grid-template-columns: 1fr;
  }
  .page-top {
    flex-direction: column;
  }
}